Curriculum Vitae - Kathy Hill


Citizenship: Dual New Zealand, USA

Employment Address (to 2012): Ecology and Evolutionary Biology, 75 N. Eagleville Rd U-3043, University of Connecticut, Storrs, CT 06269, USA

Current Position: Kathy was seriously injured in 2012 and lives in Auckland, New Zealand. For academic inquiries contact David Marshall.


Relevant work experience and future goals

After completing a MSc on the phylogeography of a New Zealand cicada species, I have continued to be employed as a field and molecular laboratory research technician for ongoing cicada phylogeny, phylogeography and behaviour studies. In Northern Hemisphere summers (May-August 2002-present) I have conducted field research on Magicicada spp, and collected information on annual North American cicada species. In the Southern Hemisphere summers (November-March 2001-present) I have been working in the field collecting cicadas for several grants to the Simon Lab, including in New Zealand for phylogeography studies, and in Australia, Fiji, South Africa, China, Argentina and the Philippines for phylogenetic studies. In addition to collecting specimens in the field I also study cicada behaviour and record cicada songs. Along with my colleages I manage databases in the lab, and am resposible for databasing specimens collected and sent to the lab, and for creating a cicada taxonomic database. I am currently authoring or co-authoring papers to describe cicada species and/or genera in New Zealand, Australia, North America, Pakistan and the Philippines. During molecular laboratory work at the University of Connecticut I have trained undergrad students in lab techniques and curation techniques and have worked with visiting researchers. In the future I hope to continue to study all aspects of Cicadidae biology and systematics.




2004-2012 – Research Technician, C. Simon lab, Ecology and Evolutionary Biology, University of Connecticut, Storrs, Connecticut, USA

2001 – Victoria University of Wellington, School of Biological Sciences lab demonstrator: Year 1 general biology.

Sept-Dec 2000 – U.S. Geological Survey, Palila Project, Hawaii. Collecting, monitoring and rearing insects related to the lifestyle of an endangered Hawaiian bird, the Palila.

Oct-Feb 1997-1999 – Manaaki Whenua Landcare Research CRI, New Zealand, Biological Control of Weeds group: Rearing introduced insects for release in NZ to eat noxious introduced plants. Testing Integrated Pest Management (IPM) schemes for the combined use of chemical and insect control agents

1999 – Lincoln University Insect Museum, curating insect collections Lincoln University Statistics and Economics departments: Lab demonstrator for Year 2 Statistics and Year 3 Economics computer labs.

1998-2000  – Lincoln University Library Student Assistant. 1999 Head Student Assistant. Circulation desk, opening and closing of library, training new student library assistants. Lincoln University computing department. Student Computing Helpdesk, fixing computer and printer problems, training new students on the university computer system. Y2K upgrade technician.




2001 - 2004  Victoria University of Wellington, New Zealand.  MSc (Ecology), April 2005

1997 - 2000  Lincoln University, New Zealand. BSc (Entomology), May 2001





Hill KBR, Marshall DC, Marathe K, Moulds MS, Lee YJ, Pham TH, Mohagan AB, Sarkar V, Price B, Duffels JP, Schouten MA, Boer AJ, Kunte K, Simon C. 2021. The molecular systematics and diversification of a taxonomically unstable group of Asian cicada tribes related to Cicadini Latreille, 1802 (Hemiptera: Cicadidae). Invertebrate Systematics 35:570-601. Email David Marshall for a reprint. A higher-resolution version of Fig. 11, which was poorly represented by the journal, can be downloaded here.

Marshall DC, Hill KBR, Cooley JR. 2018. Multimodal life-cycle variation in 13- and 17-year periodical cicadas (Hemiptera: Cicadidae: Magicicada). Journal of the Kansas Entomological Society 90(3):211-226. Issue dated July 2017, but first published online in June, 2018.


Marshall DC, Moulds MS, Hill KBR, Price BW, Wade EJ, Owen CL, Goemans G, Marathe K, Sarkar V, Cooley JR, Sanborn, AF, Kunte K, Villet MH, Simon C. 2018. A molecular phylogeny of the cicadas (Hemiptera: Cicadidae) with a review of tribe and subfamily classification. Zootaxa 4424(1):1-64.


Cooley JR, Marshall DC, Hill KBR. 2018. A specialized fungal parasite (Massospora cicadina) hijacks the sexual signals of periodical cicadas (Hemiptera: Cicadidae: Magicicada). Scientific Reports 8:1432. DOI:10.1038/s41598-018-19813-0


Marshall DC, Hill KBR. 2017. A new genus for North American Cicadetta species (Hemiptera: Cicadidae) Zootaxa 4306: 537-550.


Marshall DC, Hill KBR. 2017. A new Neotibicen cicada subspecies (Hemiptera: Cicadidae) from the southeastern USA forms hybrid zones with a widespread relative despite a divergent male calling song. Zootaxa 4272: 529-550.


Ewart A., Popple LW, Hill KBR. 2017. Five new species of grass cicadas in the genus Graminitigrina (Hemiptera: Cicadidae: Cicadettinae: Cicadettini) from Queensland and Northern Territory, Australia: comparative morphology, songs, behaviour and distributions. Zootaxa 4228: 1-62. Available here:


Owen CL, Marshall DC, Hill KBR, Simon C. 2016. How the aridification of Australia structured the biogeography and influenced the diversification of a large lineage of Australian cicadas. Systematic Biology doi: 10.1093/sysbio/syw078


Lee Y-J, Marshall DC, Mohagan A, Hill KBR. 2016. Cicadas (Hemiptera: Cicadidae) from Camiguin of Mindanao Province and Dinagat Island in the Philippines, with a new genus and three new species. Zootaxa 4097: 187-202.


Cooley JR, Kritsky G, Marshall DC, Hill KBR. 2016. A GIS-Based Map of Periodical Cicada Brood XIII in 2007, with Notes on Adjacent Populations of Broods III and X (Hemiptera: Magicicada spp.). American Entomologist 62: 241-246.


Marshall DC, Hill KBR, Moulds MS, Vanderpool D, Cooley J, Mohagan A, Simon C. 2016. Inflation of Molecular Clock Rates and Dates: Molecular Phylogenetics, Biogeography, and Diversification of a Global Cicada Radiation from Australasia (Hemiptera: Cicadidae: Cicadettini). Systematic Biology 65: 16-34.


Hill KBR, Marshall DC, Moulds MS, Simon C. 2015. Molecular phylogenetics, diversification, and systematics of Tibicen Latreille 1825 and allied cicadas of the tribe Cryptotympanini, with three new genera and emphasis on species from the USA and Canada (Hemiptera: Auchenorrhyncha: Cicadidae). Zootaxa 3985: 219-251.


Moulds MS, Hill KBR. 2015. Phylogeny for the Tribe Thophini (Cicadoidea: Cicadidae) with the Description of a New Subspecies of Thopha sessiliba Distant from Western Australia. Records of the Australian Museum 67: 55-66.


Marshall DC, Hill KBR. 2014. Comments on Tibicina Amyot, 1847 and Lyristes Horvath, 1926 (Insecta, Hemiptera, Homoptera): proposed conservation by the suppression of Tibicen Berthold, 1827 [?Latreille, 1825], and concerning the type species of Cicada Linnaeus, 1758. Bulletin of Zoological Nomenclature 71: 103-107.


Hill KBR, Marshall DC. 2013. The song, morphology, habitat, and distribution of the elusive North American cicada Okanagana viridis (Auchenorrhyncha: Cicadidae). Annals of the Entomological Society of America 106: 598-603.


Hill KBR.  2012. Chorus Cicada, Amphipsalta zelandica (Boisduval), males calling with only wing-clicks. The Weta 43: 15-20.


Marshall DC, Hill KBR, Marske KA, Chambers C, Buckley TR, Simon C. 2012. Limited, episodic diversification and contrasting phylogeography in a New Zealand cicada radiation. BMC Evolutionary Biology 12: 177.


Marshall DC, Hill KBR, Cooley JR, Simon C. 2011. Hybridization, mitochondrial DNA phylogeography, and prediction of the early stages of reproductive isolation: Lessons from New Zealand cicadas (Genus Kikihia). Systematic Biology 60: 482-502.


Cooley J, Kritsky G, Edwards MJ, Zyla JD, Marshall DC, Hill KBR, Bunker G, Neckermann M, Troutman R, Yoshimura J, Simon C. 2011. Periodical cicadas (Magicicada spp.): a GIS-based map of Broods XIV in 2008 and "XV" in 2009. American Entomologist 57: 144-150.


Marshall DC, Cooley JR, Hill KBR. 2011. Developmental plasticity of life-cycle length in thirteen-year periodical cicadas (Hemiptera: Cicadidae). Annals of the Entomological Society of America 104: 443-450.


Lee YJ, Hill KBR. 2010. Systematic revision of the genus Psithyristria Stål (Hemiptera: Cicadidae) with seven new species and a molecular phylogeny of the genus and higher taxa. Systematic Entomology 35: 277-305.

Ahmed Z, Sanborn AF, Hill KBR. 2010. A new species of the cicada genus Cicadatra from Pakistan. Zoology in the Middle East 51: 75-81.

Hill KBR, Marshall DC.  2009.  Confirmation of the cicada Tibicen pronotalis stat. nov. (= T. walkeri, Hemiptera: Cicadidae) in Florida: finding singing insects through their songs.  Zootaxa 2125:63-66

Cooley JR, Kritsky G, Zyla JD, Edwards MJ, Marshall DC, Hill KBR, Krauss R, Simon C.  2009.  The distribution of Periodical Cicada Brood X in 2004.  American Entomologist 55: 106-112

Marshall DC, Hill KBR.  2009.  Versitile aggressive mimicry of cicadas by an Australian predatory katydid. PLoS One 4: e4185. doi:10.1371/journal.pone.0004185.

Hill KBR, Simon C, Marshall DC, Chambers GK.  2009.  Surviving glacial ages within the Biotic Gap: phylogeography of the New Zealand cicada Maoricicada campbelli.  Journal of Biogeography 36:675-692.

Marshall DC, Hill KBR, Fontaine KM, Buckley TR, Simon C. 2009. Glacial refugia in a maritime temperate climate: Cicada (Kikihia subalpina) mtDNA phylogeography in New Zealand. Molecular Ecology 18: 1995-2009.

Marshall DC, Slon K, Cooley JR, Hill KBR, Simon C.  2008.  Steady Plio-Pleistocene diversification and a 2-million year sympatry threshold in a New Zealand cicada radiation.  Molecular Phylogenetics and Evolution 48: 1054-1066.

Hill KBR, Marshall DC.  2008.  Cicadas (Box spread). In: The Natural History Of Canterbury, M. Winterbourne, G. Knox, C. Burrows and I. Marsden eds. Canterbury University Press, 921pp.

Cooley JR, Marshall  DC, Hill KBR, Simon C.  2006.  Reconstructing asymmetrical reproductive character displacement in a periodical cicada contact zone.  Journal of Evolutionary Biology 19: 855-868.

Hill KBR, Marshall DC, Cooley JR.  2005.  Crossing Cook Strait: Possible human transportation and establishment of two New Zealand cicadas from North Island to South Island (Kikihia scutellaris and K. ochrina, Hemiptera: Cicadidae).  New Zealand Entomologist 28: 71-80.

Hill KBR.  2005.  Phylogeography of Maoricicada campbelli (Hemiptera: Cicadidae): mtDNA evidence interpreted using phylogenetics and nested clade analysis.  MSc Thesis, Victoria University of Wellington, Wellington, New Zealand.


Recent presentations at scientific meetings


2010. An overview of Cicadidae. Oral presentation: Arthropod Research Workshop, July 6, Central Mindanao University, Musuan, Mindanao, Philippines.

2010. Collecting, labelling and storing insects. Oral presentation: Arthropod Research Workshop, July 6, Central Mindanao University, Musuan, Mindanao, Philippines.

2010. Preliminary data on the phylogeny of the Family Cicadidae. Oral presentaion: 3rd NSF-PEET Auchenorrhyncha workshop, Apr 10-11, New York State Museum, Albany, New York, USA.

2009. Molecular phylogenetics and diversity of the Cicadidae with special reference to the tribe Cryptotympanini. Oral presentation: Seminar organised by Dr Zhongren Lei, Institute of Plant Protection, Beijing, China

2008. Tibicen phylogeny and systematics. Oral presentation: 2nd NSF-PEET Auchenorrhyncha workshop, Nov 1-2, University of Connecticut, Storrs, USA.

2008. Finding and defeating numts (NUclear copies of MiTochondrial DNA). Oral presentation: 2nd NSF-PEET Auchenorrhyncha workshop, Nov 1-2, University of Connecticut, Storrs, USA. Slideshow available at

2005. Phylogeography of a widespread New Zealand subalpine cicada, Maoricicada campbelli (Hemiptera, Cicadidae). Oral presentation: Systematics Symposium . 12th International Auchenorrhyncha Congress,  Berkeley, CA, USA, August 8-12, 2005. Abstract available at

2004    The evolution of female signalling in cicadas - a preliminary report. Oral presentation: Phylogeny and Evolution of Sternorrhyncha and Auchenorrhyncha: The First Charles P. Alexander Symposium on Insect Systematics. Amherst, MA, U.S.A.

2003. Using Nested Clade Analysis to understand the population structure of a New Zealand cicada, Maoricicada campbelli. Oral presentations: 52nd Annual Conference, Entomological Society of New Zealand, Victoria University of Wellington, 13-16 April 2003 Wellington, New Zealand; School of Biological Sciences, Victoria University of Wellington, Wellington, New Zealand 14 March 2003; Kaikoura 03: a meeting of Mathematicians and Phylogeneticists, Kaikoura, New Zealand; Poster presentation: Evolution meetings: Society for the study of evolution and Society of Systematic Biologists. Chico, California, U.S.A.

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